The Joseph cells extend from the caudal end of the anterior vesicle ( or cerebral vesicle ) to the boundary between myomeres 3 and 4, where the Hesse organs begin and continue nearly to the tail.
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The cephalic end of the neural groove exhibits several dilatations, which, when the tube is closed, assume the form of three vesicles; these constitute the three primary cerebral vesicles, and correspond respectively to the future fore-brain ( prosencephalon ), mid-brain ( mesencephalon ), and hind-brain ( rhombencephalon ).
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